guanine [guanines]◼◼◼(substance obtained from guano) noun [UK: ˈɡwɑː.ˌnin] [US: ˈɡwɑː.ˌnin]DNA is composed of four nucleotides: adenine, thymine, cytosine and guanine. = L'ADN est composé de quatre nucléotides : adénine, thymine, cytosine et guanine.